Yemen's interior ministry has announced it has arrested members of an alleged Iranian-led spy ring.

The spy ring operated in the Horn of Africa with the operations centre in Sanaa and was headed by a former member of Iran's Revolutionary Guard, it said.

Yemeni President Abdrabuh Mansur Hadi later warned Iran to "leave Yemen to mind its own affairs".

His warning echoes US claims that Iran is meddling in Yemen, which is still wrestling secessionism and rebellion.